Protons are actively transported using ATP out of the companion cell into the cell wall, creating an electrochemical gradient. These protons then re-enter the cell by facilitated diffusion through a co-transport protein in the cell membrane, which transports a molecule of sucrose with every proton transported. Sucrose enters the companion cell against the concentration gradient, increasing the concentration of sucrose in the cell. Sucrose diffuses down a concentration gradient from the companion cell into the sieve tube element through plasmodesmata.